Tim,
I think it is to do with break points
there is a breakpoint at 1650 pixels which shows the bottom link with the border, and the top link blue, therefore larger screens will show that design with the border and the blue colour.
below that breakpoint everything goes to how you want it
there is another breakpoint at 1120 pixels where the bottom link moves up closer to the top link,
I think that could be the answer but then I'm not expert by any means
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